Marked differences, however, were scored between sites under obviously different light conditions, notably in shade (closed canopy) or sun (open canopy); estimates range from 3.8 (SE $\pm$ 0.3) to 7.0 (SE $\pm$ 0.6) nodes$\cdot$ shoot$^{-1} \cdot$ year$^{-1}$, respectively (note, these were comparable with branch shoots on mature trees nearby).